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Over time the soil can compact producing a type of anerobic bacteria that will consume nitrate and produce gas that will feed a specific type of cyanobacteria. At least that is what happened in my tank. I added a little current to 'disturb' the substrate and it went away.... It took a lot of searching for me to figure out what was happening. The trick is that there are different type of anaerobic bacteria and they produce different type of by-product which has different impact on the tank.... One hint i had of what was happening is the nitrate level stayed around 1ppm despite an extremely dense fish population for several months. I guess that was the positive ;)
 
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