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"1/4 inch, shrimp-like animals get washed out and dart around until they can get back to their hornwort refuge" ... those sound like amphipods ("scuds'), from your description.
 

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They can. We had goldfish hitch a ride into our pond on plants. as soon as they were visible, I got the blame. I maintained my innocence and suggested it was eggs on the roots of the water lettuce. It was kinda cool <3
 
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