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Just a hunch: Try growing it with fewer fish and much less N and P. I have some H. polysperma in a guppy tank that is jammed with guppies (my guppy slum), and the plants look similar. Older leaves get full of holes and fall off. I know there is plenty of potassium and magnesium---the two nutrients whose deficiency might cause holes or die-back in older leaves. I think the injury is somehow related to too many fish and too 'rich' an environment.
